Clamps oriented backwards can occur after the formation of anastomosis, where two neighbouring clamps hint towards each other with their obtuse angle. After the continuation of hyphal growth of that hypha which formed the anastomosis with its tip, two facing clamps can be seen; reversely oriented clamps which hint towards each other by their overobtuse angle can also be found in the adjacent cell. Hyphae with only one cell possessing clamps and facing each other with their overobtuse angle, are formed by a backwards growing branch after forcation.

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